Biomedical Sciences
Professors
Ruben Abagyan, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Joseph Adams, PhD, Pharmacology
Kim E. Barrett, PhD, Medicine
Timothy D. Bigby, MD, Clinical Medicine
Roland C. Blantz, MD, Medicine (Emeritus)
Rolf Bodmer, PhD, Pediatrics (Adjunct)
Gerry R. Boss, MD, Medicine
Joan Heller Brown, PhD, Pharmacology (Chair)
Laurence L. Brunton, PhD, Pharmacology/Medicine (Emeritus)
Nigel Calcutt, PhD, Pathology
Dennis A. Carson, MD, Medicine (Emeritus)
Webster K. Cavenee, PhD, Medicine
Geoffrey Chang, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Ju Chen, PhD, Medicine (In Residence)
David Cheresh, PhD, Pathology
Shu Chien, MD, PhD, Bioengineering/Medicine
Mario Chojkier, MD, Medicine (In Residence)
Jerold J. M. Chun, PhD, Pharmacology (Adjunct)
Don W. Cleveland, PhD, Medicine/Cellular and Molecular Medicine (Chair)
Sara Courtneidge, PhD, Pharmacology (Adjunct)
Antonio De Maio, PhD, Surgery
Edward Dennis, PhD, Pharmacology/Chemistry and Biochemistry
Arshad Desai, PhD, Cellular and Molecular Medicine
Jack E. Dixon, PhD, Pharmacology/Cellular and Molecular Medicine/Chemistry and Biochemistry (Dean, Scientific Affairs)
Daniel Donoghue, PhD, Chemistry and Biochemistry
Steven F. Dowdy, PhD, Cellular and Molecular Medicine
Mark H. Ellisman, PhD, Neurosciences
Peter Ernst, DVM, PhD, Pathology
Jeffrey Esko, PhD, Cellular and Molecular Medicine
Ronald M. Evans, PhD, Biology (Adjunct)
Sylvia Evans, PhD, Pharmacy/Pharmacology
Marilyn G. Farquhar, PhD, Cellular and Molecular Medicine
Gen-Sheng Feng, PhD, Pathology
James R. Feramisco, PhD, Medicine/Pharmacology (Emeritus)
Susan Ferro-Novick, PhD, Cellular and Molecular Medicine
Kelly Frazer, PhD, Pediatrics
Hudson Freeze, PhD, Medicine (Adjunct)
Theodore Friedmann, MD, Pediatrics
Xiang-Dong Fu, PhD, Cellular and Molecular Medicine
Minoru Fukuda, PhD, Pathology (Adjunct)
Frank Funari, PhD, Pathology
Richard Gallo, MD, PhD, Medicine
William Gerwick, PhD, Oceanography/Skaggs School of Pharmacy and Pharmaceutical Sciences
Gordon Gill, MD, Medicine (Emeritus)
Frances Gillin, PhD, Pathology (Emeritus)
Michael Gilson, MD, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Mark Ginsberg, MD, Medicine
Christopher K. Glass, MD, PhD, Cellular and Molecular Medicine/Medicine
Joseph G. Gleeson, MD, Neurosciences
Lawrence S. B. Goldstein, PhD, Cellular and Molecular Medicine
Steven Gonias, MD, PhD, Pathology (Chair)
Kun-Liang Guan, PhD, Pharmacology
John C. Guatelli, PhD, Medicine
Gabriel Haddad, MD, Pediatrics (Chair)/Neurosciences
Bruce Hamilton, PhD, Medicine
Tracy Handel, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Dorit Hanein, PhD, Pathology (Adjunct)
Alan Hargens, PhD, Orthopaedic Surgery
Stephen Hedrick, PhD, Cellular and Molecular Medicine
Michael C. Hogan, PhD, Medicine (Adjunct)
Vivian Hook, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Stephen B. Howell, MD, Medicine
Paul A. Insel, MD, Pharmacology/Medicine
Yishi Jin, PhD, Cellular and Molecular Medicine/Biological Sciences
Martin F. Kagnoff, MD, Medicine
Mark Kamps, PhD, Pathology
Michael Karin, PhD, Pharmacology
John Kelsoe, MD, Psychiatry
Thomas J. Kipps, MD, PhD, Medicine
Richard Klemke, PhD, Pathology
Richard Kolodner, PhD, Medicine
Elizabeth A. Komives, PhD, Chemistry and Biochemistry
Albert La Spada, MD, PhD, Cellular and Molecular Medicine
Hyam L. Leffert, MD, Pharmacology
Greg Lemke, PhD, Neurosciences (Adjunct)
Fred Levine, MD, PhD, Pediatrics (Emeritus)
Willis Li, PhD, Medicine
Richard Lieber, PhD, Orthopaedics and Bioengineering
Stuart Lipton, MD, PhD, Neurosciences (Adjunct)
Marianne Manchester, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Martin Marsala, MD, Anesthesiology
James Andrew McCammon, PhD, Chemistry and Biochemistry/Pharmacology
Pamela L. Mellon, PhD, Reproductive Medicine/Neurosciences
Mark Mercola, PhD, Bioengineering
Yury Miller, MD, PhD, Medicine
Paul Mischel, MD, Pathology
Andrew Mizisin, PhD, Pathology
Marc Montminy, MD, PhD, Biological Sciences (Adjunct)
Alexandra Newton, PhD, Pharmacology
Sanjay Nigam, MD, Cellular and Molecular Medicine/Pediatrics
Victor Nizet, MD, Pediatrics and Skaggs School of Pharmacy and Pharmaceutical Sciences
Peter Novick, PhD, Cellular and Molecular Medicine
Daniel T. O’Connor, MD, Medicine
Karen Oegema, PhD, Cellular and Molecular Medicine
Jerrold M. Olefsky, MD, Medicine
Robert Oshima, PhD, Pathology (Adjunct)
Ana Pajor, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Elena Pasquale, PhD, Pathology (Adjunct)
Maurizio Pellecchia, PhD, Pathology (Adjunct)
Renate B. Pilz, MD, Medicine (In Residence)
Frank L. Powell Jr., PhD, Medicine
Morton P. Printz, PhD, Pharmacology (Emeritus)
Oswald Quehenberger, PhD, Medicine
James Quigley, PhD, Pathology (Adjunct)
Tariq Rana, PhD, Pediatrics (Adjunct)
Anjana Rao, PhD, Pharmacology (Adjunct)
Eyal Raz, MD, Medicine (In Residence)
Bing Ren, PhD, Cellular and Molecular Medicine
Tannishtha Reya, PhD, Pharmacology
Douglas D. Richman, MD, Pathology/Medicine (In Residence)
Robert Rickert, PhD, Pathology (Adjunct)
Ze’ev Ronai, PhD, Pathology (Adjunct)
Michael G. Rosenfeld, MD, Medicine
Robert Ross, MD, Medicine (In Residence)
Guy Salvesen, PhD, Pathology (Adjunct)
Maike Sander, MD, Pediatrics/Cellular and Molecular Medicine
David Schlaepfer, PhD, Reproductive Medicine
Geert Schmid-Schoenbein, PhD, Bioengineering
Nicholas Schork, PhD, Psychiatry and Biostatistics
Sanford Shattil, MD, Medicine
Shunichi Shimasaki, PhD, Reproductive Medicine
Aleem Siddiqui, PhD, Medicine
Evan Snyder, MD, PhD, Pediatrics (Associate Physician)
Deborah Spector, PhD, Cellular and Molecular Medicine
Stephen A. Spector, MD, Pediatrics
Charles F. Stevens, MD, PhD, Pharmacology (Adjunct)
Palmer W. Taylor, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences (Dean)
Susan S. Taylor, PhD, Chemistry and Biochemistry
David Traver, PhD, Cellular and Molecular Medicine
JoAnn Trejo, PhD, Pharmacology
Roger Y. Tsien, PhD, Pharmacology/Chemistry and Biochemistry
Robert H. Tukey, PhD, Pharmacology/Chemistry and Biochemistry
Ajit P. Varki, MD, Medicine
Judith A. Varner, PhD, Pathology
Francisco Villarreal, MD, PhD, Medicine (Adjunct)
Joseph Vinetz, MD, Medicine
Peter D. Wagner, MD, Medicine
Jean Wang, PhD, Medicine
Nicholas J. G. Webster, PhD, Medicine (In Residence)
Robert Wechsler-Reya, PhD, Pediatrics (In Residence)
John B. West, MD, PhD, Medicine (Emeritus)
Miles Wilkinson, PhD, Reproductive Medicine
Elizabeth Winzeler, PhD, Pediatrics
Joseph L. Witzum, MD, Medicine
Tony L. Yaksh, PhD, Anesthesiology/Pharmacology
Gene Yeo, PhD, Cellular and Molecular Medicine
Maurizio Zanetti, MD, Medicine (Emeritus)
Dong-Er Zhang, PhD, Pathology/Biological Sciences
Kang Zhang, MD, PhD, Ophthalmology
Associate Professors
Adah Almutairi, PhD, Pharmacy/Pharmaceutical Sciences
Radha Ayyagari, PhD, Radiology/Ophthalmology
Lars Bode, PhD, Pediatrics
Martina Buck, PhD, Medicine (Adjunct)
Jack Bui, MD, PhD, Pathology
Shane Crotty, PhD, Medicine (Adjunct)
Nazneen Dewji, PhD, Medicine (Adjunct)
Pieter Dorrestein, PhD, Pharmacy/Chemistry and Biochemistry
Seth Field, MD, PhD, Medicine
Pascal Gagneux, PhD, Cellular and Molecular Medicine (Adjunct)
Pradipta Ghosh, MD, Medicine
Asa Gustafsson, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Catriona Jamieson, MD, PhD, Medicine
Mark Lawson, PhD, Reproductive Medicine
Anne Murphy, PhD, Pharmacology (Adjunct)
Hemal Patel, PhD, Anesthesiology
Jonathan Sebat, PhD, Psychiatry/Cellular and Molecular Medicine
Marion Sewer, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Jianhua Shao, MD, PhD, Pediatrics
Christina Sigurdson, DVM, PhD, Pathology
Dwayne Stupack, PhD, Reproductive Medicine
Bruce Torbett, PhD, Pathology (Adjunct)
Jing Yang, PhD, Pharmacology/Pediatrics
Kun Zhang, PhD, Bioengineering
Binhai Zheng, PhD, Neurosciences
Huilin Zhou, PhD, Cellular and Molecular Medicine
Assistant Professors
Nuno Bandeira, Skaggs School of Pharmacy and Pharmaceutical Sciences
Rafael Bejar, MD, PhD, Internal Medicine
John Chang, MD, Medicine
Stephanie Cherqui, PhD, Pediatrics (In Residence)
Neil Chi, MD, PhD, Medicine
Kevin Corbett, PhD, Cellular and Molecular Medicine
Adam Engler, PhD, Bioengineering/UC San Diego Stem Cell Institute
Wenxian Fu, PhD, Pediatrics
Thomas Hnasko, PhD, Neurosciences
Mohit Hain, MD, PhD, Medicine and Pharmacology
William Joiner, PhD, Pharmacology
Alexander Kauffman, PhD, Reproductive Medicine
Jonathan Lin, MD, PhD, Pathology
Gerald P. Morris, MD, PhD, Pathology
Alysson Muotri, PhD, Pediatrics/Cellular and Molecular Medicine
Mana Parast, MD, PhD, Pathology
Ekihiro Seki, MD, PhD, Medicine (Adjunct)
George Sen, PhD, Medicine/Cellular and Molecular Medicine
Farah Sheikh, PhD, Medicine (Adjunct)
Alexey Terskikh, PhD, Pediatrics (Adjunct)
Dong Wang, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Karl Willert, PhD, Cellular and Molecular Medicine
Benjamin Yu, MD, PhD, Medicine (In Residence)